These are the special missions that are unique for each career. They are available every 5 levels, starting at level 5. You will receive them from any of the skill trainers for your chosen career - he will have a mission symbol ("!") as soon as he has to offer you one.
Career missions offer special, career specific rewards - most of them give you an additional costume option, and at levels 10, 20, 30, 40 and 50 you get the skills for your profession's Career skill line. They are also tailored for each career as far as gameplay is concerned. This is most noticeable in the Freetrader Career Missions, which often only require a minimum of combat (or none at all) and instead tend to be about negotiations and problem-solving.
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